Rank 1: Greatly improved.
All the main issues have been addressed, and I don't mind that you didn't agree with me on everything, because a lot of the things I said were subjective. The only thing I'd still suggest changing is in Level 1 -- the long first paragraph is a bit daunting for the very first text you ask the player to read, so I'd put a paragraph break after "assigning them skills".
By Rank 2, longer texts are a bit more acceptable since the player has (hopefully) gotten a taste for NeoLemmix and is more willing to spend time reading, knowing they get rewarded with cool levels. So I won't say too much about the length here, but I'll look out in case there are cuts I think you should make.
Rank title: It wouldn't be clear to a new player what you mean by "functions", and "interactables" is cumbersome (and the very first object introduced, background decoration, is
not interactive). I would just call this rank "Objects" even though it also tutorialises some hotkeys; most of the focus is on objects, and the hotkeys are just introduced along the way when they become relevant.
Level 1: It's the "paws" button, not "paw". Hence the pun.
Level 2: "Steel can't be destroyed" is true but you could make the stronger statement "Steel can't be damaged". This would also avoid the repetition of destroyed/destructive.
Let's should be
lets. "Let's" is short for "let us" and is used to make suggestions, as in "Let's go for a walk."
Level 3: No issues.
Level 4: Typo: "trun" for "turn" at the start of the second paragraph.
Level 5: Cut "to make sure he faces the correct direction".
Suggested rewrite for second paragraph: "Levels may begin with some pre-placed lemmings, which are active straight away. They may begin with permanent skills already assigned to them."
Level 6: "One-way walls" has only one hyphen. Fifth line should be "bashers, fencers
or miners".
Second paragraph: "basher" should be "bashers".
Level 7: Comma after "though". Spelling: "necessary".
This level (and also Level 9) requires builders to be placed precisely. I can't remember, did the first rank talk about skill shadows at any point? Might be good to fit in a mention somewhere, and it will be tricky to do this in rank 2 which already has longer texts.
Level 8: Comma after "64-pixel fall".
Level 9: "They only can" should be "They can only".
Level 10: "Splat pads" are not hyphenated, and "anti-splat pads" have only one. "In contrast" should be "by contrast".
Level 11: The wall of text is extremely daunting. Fortunately, I think this one needs a full rewrite anyway, because it shouldn't mention specific hotkeys or time intervals since the player may have a different hotkey setup, and that should shorten it quite a bit anyway.
This was a fun level!
Level 12: Replace second sentence with "In this case they are marked with the icon of that skill."
Suggested rewrite for second paragraph: "Normally, lemmings are equally spread among all hatches, but some hatches can be limited to spawning a certain number of lemmings." [Keep the part about exits as-is.]
The level is pretty tricky as you can't make any mistakes in where lemmings go -- though by this point we have been introduced to rewinding, so it might be okay to leave it.
Level 13: Typo: "hightlight". Pretty neat level
Level 14: "Who shall nullify" should be "to nullify". Spelling: "incoming".