Ok, got it working now. Looks like it's not so much the ROM version, as that I haven't edited the uae config file correctly to point to the correct paths for the ROM file and Hardfile. Once I re-loaded the files through the WinUAE UI instead of trying to hand-edit the .uae file, it works.
For those who wants to get it working in WinUAE:
- make sure you use a pretty recent version of
WinUAE (at least not some old version from like 7 years ago)
- after loading the .uae config file included in platon42's zip, go to the "ROM" and "Hard drive" sections of WinUAE properties and set the ROM and Hardfile to the corresponding files on your computer. Whatever was set in the .uae config file probably won't work since the filepaths were based on platon42's computer and not yours. The hardfile is the .hdf file in platon42's zip.
- I've attached KickStart ROM v3.1 here. You probably can use any KS ROM of at least v3.0.
- (optional) also go to "CPU and FPU", and for CPU Emulation Speed,
don't pick "fastest possible". I found that fastest possible does not work well performance-wise when actually running Lemmings 2, at least on my computer. The option to be closer to real speed works fine on my computer for both running the game as well as the editor.
- Save the configuration once you make the changes, of course.